There is compelling evidence that mitochondria and chloroplasts were once primitive bacterial cells. This evidence is described in the endosymbiotic theory. How did this theory get its name? Symbiosis occurs when two different species benefit from living and working together. When one organism actually lives inside the other it's called endosymbiosis. The endosymbiotic theory describes how a large host cell and ingested bacteria could easily become dependent on one another for survival, resulting in a permanent relationship. Over millions of years of evolution, mitochondria and chloroplasts have become more specialized and today they cannot live outside the cell.
It's Just a Theory
In everyday speech, people use the word theory to mean an opinion or speculation not necessarily based on facts. But in the field of science, a theory is a well established explanation based on extensive experimentation and observation. Scientific theories are developed and verified by the scientific community and are generally accepted as fact.
Mitochondria Have DNA
Mitochondria and chloroplasts have striking similarities to bacteria cells. They have their own DNA, which is separate from the DNA found in the nucleus of the cell. And both organelles use their DNA to produce many proteins and enzymes required for their function. A double membrane surrounds both mitochondria and chloroplasts, further evidence that each was ingested by a primitive host. The two organelles also reproduce like bacteria, replicating their own DNA and directing their own division.
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- Summary
Learning Objectives
- Briefly describe what is meant by the endosymbiotic theory.
- Give some evidence supporting the theory that mitochondria and chloroplasts may have arisen from prokaryotic organisms.
It is thought that life arose on earth around four billion years ago. The endosymbiotic theory states that some of the organelles in today's eukaryotic cells were once prokaryotic microbes. In this theory, the first eukaryotic cell was probably an amoeba-like cell that got nutrients by phagocytosis and contained a nucleus that formed when a piece of the cytoplasmic membrane pinched off around the chromosomes. Some of these amoeba-like organisms ingested prokaryotic cells that then survived within the organism and developed a symbiotic relationship. Mitochondria formed when bacteria capable of aerobic respiration were ingested; chloroplasts formed when photosynthetic bacteria were ingested. They eventually lost their cell wall and much of their DNA because they were not of benefit within the host cell. Mitochondria and chloroplasts cannot grow outside their host cell.
Evidence for this is based on the following:
- Chloroplasts are the same size as prokaryotic cells, divide by binary fission, and, like bacteria, have Fts proteins at their division plane. The mitochondria are the same size as prokaryotic cells, divide by binary fission, and the mitochondria of some protists have Fts homologs at their division plane.
- Mitochondria and chloroplasts have their own DNA that is circular, not linear.
- Mitochondria and chloroplasts have their own ribosomes that have 30S and 50S subunits, not 40S and 60S.
- Several more primitive eukaryotic microbes, such as Giardia and Trichomonas have a nuclear membrane but no mitochondria.
Although evidence is less convincing, it is also possible that flagella and cilia may have come from spirochetes.
Example \(\PageIndex{1}\)
- Briefly describe what is meant by the endosymbiotic theory.
- Give three points of evidence supporting the theory that mitochondria and chloroplasts may have arisen from prokaryotic organisms.
Solutions
- The endosymbiotic theory states that some of the organelles in eukaryotic cells were once prokaryotic microbes.
- Mitochondria and chloroplasts are the same size as prokaryotic cells and divide by binary fission.
- Mitochondria and chloroplasts have their own DNA which is circular, not linear.
- Mitochondria and chloroplasts have their own ribosomes which have 30S and 50S subunits, not 40S and 60S.
Summary
The endosymbiotic theory states that mitochondria and chlopoplasts in today's eukaryotic cells were once separate prokaryotic microbes.
This page titled 7.8: The Endosymbiotic Theory is shared under a CC BY 4.0 license and was authored, remixed, and/or curated by Gary Kaiser via source content that was edited to the style and standards of the LibreTexts platform; a detailed edit history is available upon request.